About the Herbarium
Millersville University’s James C. Parks Herbarium (registered among the World's herbaria as MVSC) is a regional center for biodiversity research and informatics. Its mission is the pursuit and dissemination of knowledge about biodiversity, emphasizing plant diversity, and its products include research articles, web and phone applications, an eJournal, and posters (listed below). The core facility includes a regionally focused botanical library and a collection of 23,000 preserved plant specimens (photo of the Herbarium). These specimens serve generally as
- historical records of regional species' distributions,
- the basis for new species descriptions and other taxonomic studies,
- teaching aids,
- reference specimens for applications requiring accurate species identification (e.g., forensics or taxonomic and floristic surveys)
- archived voucher specimens to document the species identity of plants used for various other biological investigations.
Adjacent facilities and equipment includes a molecular lab, greenhouses, and light & electron microscopes. Since 2004, the herbarium has been named in honor of its late curator, James C. Parks. The Herbarium seal consists of illustrations of Plowmanianthus grandifolius and P. dressleri (Neotropical spiderworts) bound by three concentric rings containing the phrases "The James Clair Parks Herbarium" and "Millersville University 1855," separated from eachother by Quercus alba (white oak) leaves.
Herbarium Web Apps
Herbarium Large-format Color Posters
Herbarium Publications
2. An asterisk (*) indicates student author.
2023
Veléz-Negrón YI, LL Simbaña-Carrera, CM Soto-Ramos, O Medina, E Dinkel, CR Hardy, LI Rivera-Vargas, L Ramos-Sepúlveda. 2023. First report of bacterial pineapple heart rot caused by Dickeya zeae in Puerto Rico. Plant Disease 107: 210. [https://doi.org/10.1094/PDIS-01-22-0174-PDN]
2018
Hardy CR, NW Hardy. 2018. Adapting traditional field activities in natural history education to an emerging paradigm in biodiversity informatics. American Biology Teacher 80: 501-519.
2017
Faden RB, CR Hardy. 2017. Commelinaceae, the dayflower and spiderwort family. In New Manual to the Vascular Plants of the Northeastern United States and Adjacent Canada. New York Botanical Garden Press, New York. [http://dx.doi.org/10.21135/893275471.047]
2016
Hardy NW, CR Hardy. 2016. NatureAtlas.org: exposing undergraduate computer science students to opportunities and applications in biodiversity informatics. 2016 World Congress on Engineering 1: 209-211 (ISBN: 978-988-19253-0-5).
O'Meara BC, SD Smith, WS Armbruster, LD Harder, CR Hardy, LC Hileman, L Hufford, A Litt, S Magallón, SA Smith, PF Stevens, CB Fenster, PK Diggle. 2016. Non-equilibrium dynamics and floral trait interactions shape extant angiosperm diversity. Proceedings of the Royal Society B 283: 20152304 (DOI: 10.1098/rspb.2015.2304).
2013
Hardy CR, CD Steinhart. 2013. Forensic botany in the resolution of an agricultural vandalism case. Journal of Forensic Identification 63: 144-154.
2012
Hardy CR. 2012. A primer on forensic microscopy. Pp 107-118 (Chapter 6) in DW Hall, JH Byrd (eds) Forensic Botany: A Practical Guide. John Wiley & Sons, Ltd. London.
Hardy CR, DS Martin*[MU '11]. 2012. Plants and wildlife forensics. Pp 145-160 (Chapter 9) in JE Huffman, JR Wallace (eds) Wildlife Forensics: Methods and Applications. Wiley-Blackwell Publishing, London.
Hardy CR, J Ryndock*[MU '09]. 2012. Floral morphology and organogenesis in Tinantia pringlei, along with a review of floral developmental variation in the spiderwort family, Commelinaceae. Botanical Review 78 (4): 416-427.
Hardy CR, JR Wallace. 2012. Algae in forensic investigations. Pp 145-173 (Chapter 9) in DW Hall, JH Byrd (eds) Forensic Botany: A Practical Guide. John Wiley & Sons, Ltd. London.
Price J, CR Hardy, KS Tefend, DW Szymanski. 2012. Solute geochemical mass-balances and mineral weathering rates in small watersheds II: Biomass nutrient uptake, more equations in more unknowns, and land use/land cover effects. Applied Geochemistry 27: 1247-1265.
2011
Warren BH, FT Bakker, DU Bellstedt, B Bytebier, R Claßen-Bockhoff, LL Dreyer, D Edwards, F Forest, C Galley, CR Hardy, HP Linder, AM Muasya, K Mummenhoff, KC Oberlander, M Quint, JE Richardson, V Savolainen, BD Schrire, T van der Niet, GA Verboom, C Yesson, JA Hawkins. 2011. Consistent phenological shifts in the making of a biodiversity hotspot: the Cape flora. BMC Evolutionary Biology 11: 39.
2010
Linder HP, CR Hardy. 2010. A generic classification of the Restioneae (Restionaceae), southern Africa. Bothalia 40: 1-35.
2009
Hardy CR, LL Sloat*[MU '06], RB Faden. 2009. Floral organogenesis and the developmental basis for pollinator deception in the Asiatic dayflower Commelina communis (Commelinaceae). American Journal of Botany 96: 1236-1244.
2008
Hardy CR, NW Hardy. 2008. Simple biodiversity mashups for non-tech-savvy biologists: a demonstration using the liana flora of Pennsylvania, USA. Journal of the Torrey Botanical Society 135: 585-594.
Hardy CR, P Moline, HP Linder. 2008. A Phylogeny for the African Restionaceae, and New Perspectives on Morphology's Role in Generating Complete Species Phylogenies for Large Clades. International Journal of Plant Sciences 169: 377-390.
(click here for supporting electronic appendices, matrices, and cladograms)
2007
Hardy CR, HP Linder. 2007. Phylogeny and historical ecology of Rhodocoma (Restionaceae) from the Cape Floristic Region. Aliso 23: 213-226.
(pdf
available)
Parks JC (adapted). 2007. Lycopodiaceae: clubmass family. Pp 63-68 in Rhoads AF, TA Block (eds) The plants of Pennsylvania: an illustrated manual, 2nd ed.
Parks JC, JD Montgomery (adapted). 2007. Polypodiaceae s.l. (Fern Family). Pp 78-105 in Rhoads AF, TA Block (eds) The plants of Pennsylvania: an illustrated manual, 2nd ed.
2006
Hardy CR. 2006.
Reconstructing ancestral ecologies: challenges and
possible solutions. Diversity & Distributions 12: 7-19.
Davis JI, G Petersen, O Seberg, DW Stevenson, CR Hardy, MP Simmons, FA Michelangeli, DH Goldman, LM Campbell, CD Specht, JI Cohen. 2006. Are mitochondrial genes useful for the analysis of monocot relationships? Taxon 55: 857-870.
Petersen G, O Seberg, JI
Davis, D Goldman, DW Stevenson, LM Campbell, FA
Michelangeli, CD Specht, MW Chase, MF Fay, JC Pires, JV
Freudenstein, CR Hardy, MP Simmons. 2006.
Mitochondrial data in monocot phylogenetics. Aliso 22: 52-62.
Hardy CR, HP Linder. 2005.
Intraspecific variability and timing in ancestral
ecology reconstruction: A test case from the Cape flora. Systematic Biology 54 (2): 299-316.
Linder HP, CR Hardy. 2005.
Speciation in the
Bakker, L. W. Chatrou, B. Gravendeel and P. B. Pelser, eds. Plant species-level systematics: new perspectives on
pattern & process. Koeltz, Königstein
Linder HP, CR Hardy, F
Rutschmann. 2005. Taxon sampling effects in molecular
clock dating: An example from the African Restionaceae. Molecular Phylogenetics & Evolution 35: 569-582.
Hardy CR, RB Faden. 2004. Commelinaceae. Pp 425-427 in Smith N, S
Mori, A Henderson, DW Stevenson, SV Heald (eds) Flowering Plants of the Neotropics.
Hardy CR, RB Faden. 2004. Plowmanianthus, a new genus of Commelinaceae
with five new species from tropical America. Systematic Botany 29 (2): 316-333.
Hardy
CR, JI Davis, DW Stevenson. 2004. Floral
organogenesis in Plowmanianthus (Commelinaceae).
International Journal of Plant Sciences 165 (4):
511-519.
Davis JI, DW Stevenson, G Petersen,
O Seberg, LM Campbell, JV Freudenstein, DH Goldman, CR Hardy, FA Michelangeli, MP Simmons, CD Specht, F Vergara-Silva, MA Gandolfo. 2004. A phylogeny of the monocots, as inferred from rbcL and atpA sequence variation, and a comparison of methods for
calculating jackknife and bootstrap values. Systematic
Botany 29 (3): 467-510.
Linder HP, CR Hardy. 2004. Evolution of the species-rich
2003
Hardy
CR. 2003.
Hardy
CR, DL
Lipscomb, KC Nixon, H Ochoterena. 2003. WinClada ver. 1.0, a basic users manual.
Available upon request from Kevin C. Nixon, Bailey Hortorium, 462 Mann Library, Cornell
University, Ithaca, NY 14853.
2001
Hardy CR. 2001. Systematics of Cochliostema, Geogenanthus, and an undescribed genus in the spiderwort
family Commelinaceae. Ph.D. Dissertation,
2000
Dyer AF, JC Parks, S Lindsay. 2000. Historical review of the uncertain taxonomic status of Cystopteris dickieana R.Sim (Dickie's bladder fern). Edinburgh Journal of Botany 57 (1): 71-81.
Hardy CR,
DW Stevenson. 2000. Floral organogenesis in some species
of Tradescantia and Callisia (Commelinaceae). International Journal of Plant
Sciences 161 (4): 551-562.
Hardy
CR, DW Stevenson. 2000. Development of the
flower, gametophytes, and floral vasculature in Cochliostema odoratissimum (Commelinaceae). Botanical Journal of the Linnean Society 134
(1): 131-157.
Hardy CR, DW Stevenson, HG Kiss. 2000. Development of the flower, gametophytes, and floral
vasculature in Dichorisandra thyrsiflora (Commelinaceae). American Journal of Botany 87 (9): 1228-1239.
Parks JC, AF Dyter, S Lindsay. 2000. Allozyme, spore and frond variation in some Scottish populations of the ferns Cystopteris dickieana and Cystopteris fragilis. Edinburgh Journal of Botany 57 (1): 83-105.
Parks JC. 2000. Lycopodiaceae. Pp 64-70 in Rhoads AF, TA Block (eds) The plants of Pennsylvania: an illustrated manual.
Parks JC, JD Montgomery. 2000. Ferns. Pp 71-106 in Rhoads AF, TA Block (eds) The plants of Pennsylvania: an illustrated manual.
Stevenson DW, JI Davis, JV Freudenstein, CR Hardy, MP
Simmons, CD Specht. 2000. A phylogenetic analysis of the
monocotyledons based on morphological and molecular
character sets, with comments on the placement of Acorus and Hydatellaceae. Pp 17-24 in Wilson KL and DA Morrison (eds.) Monocots: systematics and
evolution. CSIRO Publishing:
1998
Parks JC, KA Mosser. 1998. A morphological and allozymal study of a Cystoperis tennesseensis (Dryopteridaceae) population: first report of the species in Lancaster County, PA. Journal of the Pennsylvania Academy of Sciences 71 (2): 78-83.
1989
Parks JC. 1989. Distribution of gametophytic populations of Vittaria and Trichomanes in Pennsylvania. Rhodora 91: 201-206.
1985
Parks JC. 1985. Erigenia bulbosa in Lancaster County, Pennsylvania. Bartonia 51.
Stober SS, JC Parks. 1985. Chromosomal studies of Physalis virginiana var. subglabrata and Physalis heterophylla. Bartonia 51: 65-68.
1984
Parks JC. 1984. A first report of the fern genus Vittaria in New York. Rhodora 86 (848): 421-423.
Parks JC. 1984. New Pennsylvania locality of Cryptogramma stelleri. Bartonia 50: 63.
1983
Parks JC. 1983. The fern genera Vittaria and Trichomanes in the Northeastern United States. Rhodora 85 (841): 83-92.
1973
Parks JC. 1973. A revision of North American and Caribbean Melanthera (Compositae). Rhodora 75: 169-210.
1969
Parks JC. 1969. Simple technique for demonstrating chromosomal coils using Tradescantia pollen mother cells. Proceedings of the Pennsylvania Academy of Science 43: 61-63.
1968
Parks JC. 1968. A revision of the genus Melanthera Rohr (Compositae): in North America and the Caribbean. Ph.D. Dissertation,
Vanderbilt